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uppies separated from their littermates too early often don't develop appropriate "social skills," such as learning how to send and receive signals, what an "inhibited bite" means, how far to go in play wrestling and so forth. Play is important to help puppies increase their physical coordination, social skills and learning limits. Interacting with their mother and littermates helps them learn "how to be a dog" and is also a way to explore ranking ("who's in charge").
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